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Evenhuis Article number: 18.3.51ACopyright Palaeontological Association, October 2015 Author biographies Plain-language and multi-lingual abstracts PDF version Submission: 24 June 2015. is the first of the very speciose genus Anthrax and only the second bombyliid to be described from Dominican amber. Eoanomala melas contains a number of character states that are not found together in any extant genera and, although it may fit within the tribe Villini, it does not appear to be closely related to the extant constituents of that clade. Adult bombyliids are important pollinators and many have evolved a long proboscis to facilitate nectar feeding.

is described from the Middle Eocene oil shales of the Kishenehn Formation. Department of Paleobiology, National Museum of Natural History MRC 121, Smithsonian Institution, 10th & Constitution Ave. C., 20013-7012, USA; Keywords: Bombyliidae; Anthrax; Dominican amber; Kishenehn Formation; new species; new genus Final citation: Greenwalt, Dale E., Wingerath, Jonathan G., and Evenhuis, Neal L. Two new and disparate fossil bee flies (Bombyliidae: Anthracinae) from the Americas and reassessment of 18.3.51A: 1-10palaeo-electronica.org/content/2015/1350-two-new-fossil-bee-flies Bee flies (Diptera: Bombyliidae) constitute a large and diverse family of 15 different extant subfamilies, more than 230 genera and nearly 5,000 species that, as adults, feed on nectar and/or pollen (females are obligate pollen feeders) and are found in all continents except Antarctica (Evenhuis and Greathead, 2003; Pape et al., 2011).

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A major evolutionary advance in this behavior occurred with the development of the “sand chamber”, a structure used by the female to coat eggs with particles of ground substrate (e.g., sand) prior to in-flight oviposition near potential hosts (Yeates and Greathead, 1997) to protect them from desiccation. is restricted to the Cenozoic; the oldest fossil, Paleolomatia menatensi s, is from the sediments of a Paleocene maar near Menat, France (Nel, 2008).

(2011) place the subfamily Heterotropinae basal to Mythicomyiidae.

Acceptance: 6 October 2015 In this study, two new species of anthracine bee flies are described and one previously described species is reassessed. The previously described Anthrax dentoni Lewis, 1969 was re-examined and reassigned to Anthracinae incertae sedis due to lack of morphological detail required for generic assignment. While there is uncertainty about some of the supergeneric relationships among species of this clade, Mythicomyiidae (previously Mythicomyiinae) has been shown to be basal to all Bombyliidae, and the molecular data of Trautwein et al.

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